Tuesday, December 22, 2015

[Herpetology • 2015] Cyrtodactylus petani • The fourth Bent-toed Gecko of the Genus Cyrtodactylus (Squamata: Gekkonidae) from Java, Indonesia


Farmer’s Bent-toed Gecko  | Cyrtodactylus petani 
Riyanto, Grismer & Wood, 2015

FIGURE 2. Living specimens of Cyrtodactylus petani sp. nov. from Jeladri, Pasuruan, Java.
(A) Adult male paratype MZB.Lace 11712 (SVL=56.6 mm). (B) Gravid female paratype MZB.Lace 11711. (SVL=53.8 mm).
FIGURE 6. Microhabitats where specimens of Cyrtodactylus petani sp. nov. were collected.
 (A) paddy field embankment with vegetation. (B) Rocks in the riverbank.

Abstract
Cyrtodactylus petani sp. nov. is a new species of Bent-toed Gecko from Java, Indonesia that had been masquerading under the name C. fumosus (Müller, 1895). The new species is differentiated from C. fumosus and all its Sundaland congeners by having the following combination of morphological characters: a maximum SVL of 57.2 mm; nine or ten supralabials; seven or eight infralabials; strongly tuberculate body and limbs; 20–25 paravertebral tubercles; 30–35 ventral scales; enlarged precloacal scales; enlarged femoral scales; 17–18 subdigital lamellae on the fourth toe; 31–35 continuous precloacal and femoral pores in males, pores absent in females; no precloacal groove; no enlarged median subcaudals; tubercles on anterior portion of tail; no reticulated pattern on top of head; a blotched dorsal pattern; and no paired, dark, semi-lunar shaped blotches on the nape.


Keywords: Reptilia, Cyrtodactylus, new species, Java, taxonomy, Gekkonidae



Systematics 
Cyrtodactylus petani sp. nov. 
(Cicak Jari Lengkung Petani: Farmer’s Bent-toed Gecko)


Etymology. The specific epithet petani refers to the fact that the type series was collected on a farm. Petani means a farmer in the Indonesian Language and is here treated as a noun in apposition.


FIGURE 5. Distribution map of Javan Cyrtodactylus.
Red circle is holotype of Cyrtodactylus petani sp. nov. Green circles are paratype localities of C. petani.Yellow circles are known localities of C. semiadii. The distribution of the widespread species, C. marmoratus and C. fumosus are not shown.

Discussion: 
Cyrtodactylus petani sp. nov. is the fourth species of Cyrtodactylus from Java that was not reported in previous herpetofaunal surveys of East Java. As suggested by Riyanto et al. (2014), C. fumosus from Java includes multiple undescribed species. Together, with other recent findings, including the discovery of a new Cyrtodactylus (Riyanto et al. 2014), a new Chiromanthis (Riyanto & Kurniati, 2014), a new Eutropis (Mausfeld & Böhme 2002), a suspected new species of Ptychozoon (Brown et al. 2012), a new record for Polypedates otilophus (Riyanto et al. 2009), and a relatively new Dendrelaphis (Rooijen & Vogel 2008), this discovery suggests much is yet to be learned concerning the herpetofaunal composition of Java. This discovery also indicates that the diversity of Cyrtodactylus continues to rise even in highly disturbed areas. 


Riyanto, Awal, L. L. Grismer & Wood, P. L., Jr. 2015. The fourth Bent-toed Gecko of the Genus Cyrtodactylus (Squamata: Gekkonidae) from Java, Indonesia.
Zootaxa. 
4059(2): 351–363.  DOI: 10.11646/zootaxa.4059.2.6